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NEW YORK, NY, June 12, 2008 — Invitrogen Corporation and Applera Corporation today announced a definitive merger agreement under which Invitrogen will acquire all of the outstanding shares of Applera’s Applied Biosystems Group in a cash and stock transaction valued at $6.7 billion. Covington & Burling LLP represented Greenhill & Co., which provided a fairness opinion to Applera’s Board. Invitrogen provides products and services that support academic and government research institutions and pharmaceutical and biotech companies worldwide in their efforts to improve the human condition. Applera’s Applied Biosystems serves the life science industry and research community by developing and marketing instrument-based systems, consumables, software, and services. Greenhill & Co. is an investment bank that provides financial advice on mergers, acquisitions and restructurings, and manages merchant banking funds. The Covington corporate team was led by partners Scott Smith and Jack Bodner, and associate Donald Saelinger. All three are resident in the firm’s New York office.